Percival Aloysius "Knuckles" O'Toole

Few names in the Golden Age roll off the tongue quite like Percival Aloysius "Knuckles" O'Toole — a character with a moniker that practically tells its own story. Debuting in Captain America Comics #5 in 1941, brought to life by a young Stan Lee and Charles Nicholas, Knuckles stepped onto the page during comics' earliest, most electric era, sharing those wartime adventures with none other than Captain America and Bucky Barnes themselves. His appearances are rare — a handful of catalog entries spanning an impressive stretch of publishing history from 1941 to 1990 — which makes each one a genuine treasure for Golden Age enthusiasts and completionists alike. If you love hunting down the overlooked corners of Marvel's storied origins, Knuckles O'Toole is exactly the kind of character who rewards the dig.